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Figure 5. The combination of recording the stimulus evoked CAP (triangle) with actual time recordings of lactate (black line) permit for a fuller image of the cellular interactions to emerge. At the onset of aglycemia the lactate falls virtually instantly adopted by the CAP. Glucose is taken up by astrocytes and either stored as glycogen or straight processed glycolytically to lactate. The lactate is then transported to the axons for oxidative metabolism. Removing glucose from the aCSF or inhibiting glycogen metabolism with DAB or isofagomine causes the lactate to quickly fall to zero adopted by the CAP. The conclusion from this optic nerve knowledge is that astrocytes contain glycogen and release glycogen derived lactate into the interstitial area via the MCT1, from where it's taken up into axons via the MCT2 transporter. Along with this lactate launch underneath conditions of aglycemia, where the latency to CAP failure might be accelerated by blocking lactate uptake or inhibiting glycogen metabolism, lactate can be launched tonically under baseline situations.

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